Species New to Science

Pristimantis zorro Rivera-Correa & Daza, 2020

DOI: 10.11646/zootaxa.4838.1.4

Abstract

Continuous sampling in well studied areas may lead to new amphibian species discoveries, because population dynamics allow rare species to go unnoticed for years. Based on recent sampling of frogs in the northeastern region of Colombia, here we provide genetic, morphological, and bioacoustics evidence to support the description of a new species of the genus Pristimantis. Phylogenetic analysis suggests the new species is a member of the P. lacrimosus species group, a clade of 25 species that is highly diverse in Ecuador and Peru. This new species is more closely related to allopatric species from Peru and Guyana (i.e. P. olivaceus, P. pluvialis, P. pulchridormientes, and an undetermined Pristimantis). The new species has green-yellow coloration and a remarkable brown interocular band, which is unusual in the genus and which inspired the epithet. Pristimantis zorro sp. nov. has only been found at one locality at 1860 m s.a.l., on the eastern flank of the northern Cordillera Central in Department of Antioquia, Colombia. The new species inhabits the ecotone between the humid cloud forest and open areas. Repeated visits to the type locality suggest high variation in population abundance and/or species detection, because we have occasionally observed many individuals in briefs periods of times. In addition, we discuss the biogeographical and systematic implications of our species discovery.

Keywords: Amphibia, Advertisement call, Biodiversity, Morphology, Molecular phylogenetic, Taxonomy, Systematics

Adult males of Pristimantis zorro sp. nov. in life:
(A) MHUA-A 8813, SVL 20.1 mm, holotype; (B) MHUA-A 8814, SVL 20.5 mm, paratype; (C) MHUA-A 8815, SVL 20.3 mm, paratype, adult male; (D) MHUA-A 8816, SVL 19.5 mm, paratype; (E) MHUA-A 8817, SVL 21.5 mm, paratype; (F) MHUA-A 11165, SVL 20.1 mm, paratype; (G) MHUA-A 11166, SVL 21.3 mm, paratype; (H) MHUA-A 11167, SVL 20.8 mm, paratype. Photos by Adriana Restrepo, Carlos Marín and Juan M. Daza.

Pristimantis zorro sp. nov.

Etymology. The specific name is a patronym in reference to the character El Zorro (fox in Spanish). El Zorro is the secret identity of Don Diego de la Vega, a fictional hero created in 1919 by pulp writer Johnston McCulley. The character has a distinctive black garb, coat, hat and a mask that covers the top of the head from eye level upwards. The name alludes to the facial mask of the new species.

Pristimantis olivaceus ZFMK 67132, SVL 19.7 mm, paratype.

Photo: Jörn Köhler.

Mauricio Rivera-Correa and Juan M. Daza. 2020. Out of the Blue: A New Rain Frog Species of the Genus Pristimantis (Anura: Craugastoridae) from the northern Cordillera Central in Colombia. Zootaxa. 4838(1); 83–101. DOI: 10.11646/zootaxa.4838.1.4

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twitter.com/biodiversitario/status/1298262780651347974

Resumen. El muestreo continuo en áreas bien estudiadas puede conducir a nuevos descubrimientos de especies de anfibios, debido a que la dinámica de las poblaciones permite que especies raras pasen desapercibidas durante años. Basado en unos muestreos recientes de ranas en la región nororiental de Colombia, aquí proporcionamos evidencia genética, morfológica y bioacústica que soportan la descripción de una nueva especie del género Pristimantis. El análisis filogenético sugiere que la nueva especie es miembro del grupo P. lacrimosus, un clado de 25 especies más diversas en Ecuador y Perú. Esta nueva especie está más estrechamente relacionada con especies alopátricas de Perú y Guyana (i.e. P. olivaceus, P. pluvialis, P. pulchridormientes y un Pristimantis no determinado). La nueva especie tiene una coloración verde-amarillo y una notable banda interocular marrón, que es inusual en el género y que inspiró el nombre de la nueva especie. Pristimantis zorro sp. nov. solo se ha encontrado en una localidad a 1860 m s.n.m., en el flanco oriental al norte de la Cordillera Central en el departamento de Antioquia, Colombia. La nueva especie habita el ecotono entre el bosque nublado húmedo y las áreas abiertas. Las visitas repetidas a la localidad tipo sugieren una gran variación en la abundancia de esta población y/o la detección de la especie, debido a que hemos observado individuos ocasionalmente y por breves períodos de tiempo. Adicionalmente discutimos las implicaciones biogeográficas y sistemáticas del descubrimiento de esta nueva especie.

Palabras clave.Canto de anuncio, Biodiversidad, Morfología, Filogenética molecular, Taxonomía y Sistemática

Cyrtodactylus amphiptraeus

Chomdej, Suwannapoom, Pawangkhanant, Pradit, Nazarov, Grismer & Poyarkov, 2020

ตุ๊กกายตาก || DOI: 10.11646/zootaxa.4838.2.2

Photos by C. Suwannapoom (A–B), N.A. Poyarkov (C–D), and P. Pawangkhanant (E–F).

Abstract

A new species of Cyrtodactylus from Tak Province, Thailand, Cyrtodactylus amphipetraeus sp. nov., is described using an integrative taxonomic analysis based on morphology, color pattern, and the mitochondrial gene NADH dehydrogenase subunit 2 (ND2). The phylogenetic analyses place the new species within the C. sinyineensis group which was previously thought to be endemic to the Salween Basin in southern Myanmar. The phylogeny also places C. inthanon in the C. sinyneensis group which is expanded herein to also include the group’s sister species C. doisuthep. Along with C. amphipetraeus sp. nov., these are the first three species of the C. sinyineensis group to be found outside of Myanmar east of the Tenasserim Mountains. The Tenasserim Mountain region is discussed as an area of cladogeneic turnover.

Keywords: Reptilia, Integrative taxonomy, Indochina, Tenasserim Mountains, Bent-toed gecko

Variation in dorsal pattern in Cyrtodactylus amphiptraeussp. nov. in life, from ... Pop Pra District, Tak Province, Thailand.
A. Adult holotype male (AUP-00696). B. Adult paratype male (AUP-00698).
C. General view of the adult paratype female (ZMMU R-16626). D. Close-up of the head of the same specimen showing green iris.
E. Adult male (not collected) hiding in crevice F. Juvenile specimen (not collected).

Photos by C. Suwannapoom (A–B), N.A. Poyarkov (C–D), and P. Pawangkhanant (E–F).

Cyrtodactylus amphipetraeus sp. nov.

Tak Bent-toed Gecko - ตุ๊กกายตาก

Distribution. Cyrtodactylus amphipetraeus sp. nov. is known only from the type locality at ... Cave, Mae Sot District, environs of Mae Sot, Tak Province, western Thailand; and ... Waterfall located approximately 4 km to the southwest from the cave (Fig. 1).

Etymology. The specific epithet amphipetraeus is a Latinized adjective in nominative singular, derived from Greek amphi or ἀμφί (meaning of both kinds) and petra or πέτρα (for rock). The species name is given in reference to the remarkable natural history of this species which inhabits both limestone and granite rocks. The recommended vernacular name in English is Tak Bent-toed Gecko; in Thai is Tuk kai tak - ตุ๊กกายตาก.

Siriwadee Chomdej, Chatmongkon Suwannapoom, Parinya Pawangkhanant, Waranee Pradit, Roman A. Nazarov, L. Lee Grismer and Nikolay A. Poyarkov. 2020. A New Species Cyrtodactylus Gray (Squamata: Gekkonidae) from western Thailand and the Phylogenetic Placement of C. inthanon and C. doisuthep. Zootaxa. 4838(2); 179–209. DOI: 10.11646/zootaxa.4838.2.2

Impatiens dindigulensis Ramas. Anjana & Chandra

in Ramasubbu, Anjana & Prabha, 2020.

taiwania.ntu.edu.tw

Abstract

Impatiens dindigulensis sp. nov., a new species of Balsaminaceae is described and illustrated from Kodaikkanal Wildlife Sanctuary, the Western Ghats of Tamil Nadu, India. The detailed description of taxon along with diagnostic characters between closely allied species, pollen morphology, illustration and colour photographs are provided.

Keyword: Impatiens dindigulensis, I. tanyae, I. tomentosa, taxonomy, Western Ghats

Fig. 2. Impatiens dindigulensis Ramas. Anjana and Chandra, sp. nov.
A. Habit; B. Leaves, abaxial view; C. Leaves, adaxial view; D. Pair of stipular glands; E. Flower front view; F. Flower, lateral view; G. Dorsal petal; H. Lateral united petal; I. Fruit.

Fig. 1. Impatiens dindigulensis Ramas., Anjana and Chandra, sp. nov.
A. Habit; B. Leaves, abaxial view; C. Leaves, adaxial view; D. Pair of stipular glands; E. Leaves; margin with spines; F. Stem with hairs; G. Flower, lateral view; H. Flower basal view; I. Flower front view; J. Dorsal petal. K. Lateral united petal; L. Lateral view of lower sepal; M. SEM view of Pollen; N. Fruit; O. Seeds.

Impatiens dindigulensis Ramas., Anjana & Chandra, sp. nov.

Etymology: The species is named after Dindigul district (Tamil Nadu) where the type locality Kodaikanal Wildlife Sanctury is located.

Ecology: This herb is growing under shady areas of Semi-evergreen forests with altitude ranging from 1931– 1949 msl.

Distribution: This taxon has been collected only from the Kodaikkanal Wildlife Sanctuary of the Western Ghats, Tamil Nadu, India with altitude ranging from 1931–1949 msl.

Raju Ramasubbu, Surendran Anjana and Ayyathurai Chandra Prabha. 2020. A New Species Impatiens L. (Balsaminaceae) from Kodaikkanal Wildlife sanctuary, India. Taiwania. 65(4); 426-430.

taiwania.ntu.edu.tw/abstract.php?type=abstract&id=1704

Betta nuluhon Kamal, Tan & Ng, 2020

DOI: 10.11646/zootaxa.4819.1.11

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Abstract

Betta nuluhon, new species, is described from a hill stream habitat in western Sabah. This species is allied to both B. chini and B. balunga, and differs from rest of its congeners in the B. akarensis group in having the following combination of characters: yellow iris when live; mature males with greenish-blue iridescence on opercle when live; mature fish with distinct transverse bars on caudal fin; slender body (body depth 22.1–25.2 % SL); belly area with faint reticulate pattern (scales posteriorly rimmed with black); absence of tiny black spots on anal fin; lateral scales 29–31 (mode 30); predorsal scales 20–21 (mode 20). Notes on a fresh series of B. chini are also provided.

Keywords: Taxonomy, biodiversity, Southeast Asia, Anabantiformes, riparian habitat, Pisces

FIGURE 1. Betta nuluhon, new species, ca. 50 mm SL, freshly caught male specimen, not preserved.

FIGURE 3. Betta nuluhon, new species, ZRC 61247, holotype, 62.6 mm SL.

Betta nuluhon, new species

Etymology. This new species is named for the Kadazandusun vernacular nuluhon, meaning hill. This pertains to its hill stream habitat. Used as a noun in apposition.

N. S. S. Kamal, H. H. Tan and Casey K. C. Ng. 2020. Betta nuluhon, A New Species of Fighting Fish from western Sabah, Malaysia (Teleostei: Osphronemidae). Zootaxa. 4819(1); 187–194. DOI: 10.11646/zootaxa.4819.1.11

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Cnemaspis rishivalleyensis & Hemidactylus rishivalleyensis

Agarwal, Thackeray & Khandekar, 2020.

DOI: 10.11646/zootaxa.4838.4.1

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Abstract

Two new species of geckos from the genera Cnemaspis and Hemidactylus are described from a granite outcrop in the Rishi Valley, Chittoor District, Andhra Pradesh, India. Cnemaspis rishivalleyensis sp. nov. and Hemidactylus rishivalleyensis sp. nov. are presently known only from their type locality and are 8.7 % and 10.9 % divergent from their closest known sister species, respectively. The new species are allied to the mysoriensis and murrayi clades of South Asian Cnemaspis and the brookii group of Indian Hemidactylus, respectively. The two new species can be diagnosed from regional congeners by the number and arrangement of femoral and precloacal pores and poreless scales separating these series, body size, the number of tubercles in paravertebral rows, the number of enlarged tubercles around mid-body, the number of ventral scales across mid-body, the number of lamellae beneath digit IV of pes and manus, and subtle differences in colour pattern. Four endemic geckos now have their type localities within 10 km of each other, Cnemaspis graniticola and Cyrtodactylus rishivalleyensis from Horsley Hills and the two new species from Rishi valley. The two new species are also the third endemic species each of Cnemaspis and Hemidactylus from Andhra Pradesh.

Keywords: Reptilia, Cnemaspis mysoriensis, Cnemaspis otai, Eastern Ghats, Hemidactylus treutleri, Mysore Plateau, peninsular India, taxonomy

Cnemaspis rishivalleyensis sp. nov. female, (paratype, NCBSBH724).
Hemidactylus rishivalleyensis sp. nov. female, (paratype, NCBS-BH729).

Cnemaspis rishivalleyensis sp. nov.

Cnemaspis mysoriensis [part] Giri et al. 2009a

Etymology. The specific epithet is a toponym for the Rishi Valley, the type and only known locality for the new species.

Suggested Common Name. Rishi Valley dwarf gecko

Hemidactylus rishivalleyensis sp. nov.

Hemidactylus treutleri, [part] Sreekar et al. 2010; Lajmi et al. 2016

Etymology. The specific epithet is a toponym for the Rishi Valley, the type locality for the new species. Suggested Common Name. Rishi Valley rock gecko

Ishan Agarwal, Tejas Thackeray and Akshay Khandekar. 2020. Geckos in the Granite: Two New Geckos (Squamata: Gekkonidae) from Rocky, Scrub Habitats in Rishi Valley, Andhra Pradesh, India. Zootaxa. 4838(4); 451–474. DOI: 10.11646/zootaxa.4838.4.1

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Turleania rubriguttatus Komai, 2020

DOI: 10.11646/zootaxa.4834.1.6

Abstract

A new species of the pagurid hermit crab genus Turleania McLaughlin, 1997, Turleania rubriguttatus, is described on the basis of two specimens, including one male and one female, from shallow subtidal waters in Kochi Prefecture, Japan. The new species appears close to T. albatrossae (McLaughlin & Haig, 1996), known from the Philippines, but the proximally unarmed dorsal surface of the right chela palm and the lack of a dorsomesial row of spines on the left cheliped carpus easily distinguish T. rubriguttatus n. sp. from T. albatrossae. Examination of the type material of T. similis Komai, 1999 and T. spinimanus Komai, 1999, and supplemental material from Japan, confirms that the two taxa are synonymous with T. senticosa (McLaughlin & Haig, 1996), as was suggested by previous authors. Re-examination clarified that in T. senticosa the maxilliped 3 has no developed arthrobranchs, and this led the author to assess the status of T. sinensis Han, Sha & An, 2016, which is also synonymised with T. senticosa.

Keywords: Decapoda, Kochi Prefecture, synonym, Turleania sinensis, Turleania spinimanus

Tomoyuki Komai. 2020. Turleania rubriguttatus, A New Species of Pagurid Hermit Crab (Decapoda: Anomura: Paguroidea) from Shallow Water in Japan, with Notes on T. senticosa (McLaughlin & Haig, 1996). Zootaxa. 4834(1); 96–106. DOI: 10.11646/zootaxa.4834.1.6

Deinostigma fasciculatum W.H.Chen & Y.M.Shui

in Shui, Wu, Yu, ... et Chen, 2020.

簇花奇柱苣苔 || DOI: 10.3897/phytokeys.157.32683

Abstract

A new species of Deinostigma (Gesneriaceae) from Yunnan, China, Deinostigma fasciculatum W.H.Chen & Y.M.Shui, sp. nov., has been discovered and described. In the genus, the new species is similar to D. cicatricosum (W.T. Wang) D.J. Middleton & Mich. Möller and D. cyrtocarpum(D. Fang & L. Zeng) Mich. Möller & H.J. Atkins in dark purple flowers and falcate fruit, but differs from them mainly in the inflorescences with fasciculate flowers, calyx lobes (reflexed, narrowly lanceolate and 1.2–1.3 cm long), corolla tubes (sharply contracted below middle and white outside and below throat). The above three species grow nearby non-limestone wet cliffs and geographically isolated with different distributions (the new species in Southeast Yunnan, D. cicatricosum in Eastern Guangxi and D. cyrtocarpum in Southern Guangxi and Guangdong, China).

Keywords: Deinostigma cicatricosum, Deinostigma cyrtocarpum, new species, Sino-Vietnamese border, Yunnan

Figure 1. The distribution of Deinostigma fasciculatum W.H.Chen & Y.M.Shui, sp. nov. (■),
with D. cicatricosum (W.T.Wang) D.J.Middleton & Mich.Möller (●)
and D. cyrtocarpum (D.Fang & L.Zeng) Mich.Möller & H.J.Atkins (▲).

Figure 2. Deinostigma fasciculatum W.H.Chen & Y.M.Shui, sp. nov.
A habit B mature fruits C frontal view of flower D leaf abaxial side E top and back view of flowers F top view of opened corolla showing the interior surface of corolla tube, stamens and staminodes G pistil and calyx, arrow showing the calyx H ovary, calyx and bract. (b = bract, c = calyx, d = disc).

Deinostigma fasciculatum W.H.Chen & Y.M.Shui, sp. nov.

Diagnosis: The new species is similar to D. cicatricosum and D. cyrtocarpum in dark purple flowers and falcate fruit, but differs from the latter two species in the inflorescences with fasciculate flowers (vs. with remote flowers), calyx lobes reflexed (vs. compacted), corolla tubes white outside and below throat (vs. purple) (Figs 2C, 3). The new species differs from D. cicatricosum in calyx lobes narrowly lanceolate (vs. narrowly oblong) and 1.2–1.3 cm long (vs. 0.8–1.0 cm), corolla tube sharply contracted below middle (vs. slightly contracted), capsule narrowly oblong (vs. linear) 2–2.5 cm long (vs. 3–4 cm long). It differs from D. cyrtocarpum in calyx lobes 1.2–1.3 cm long (vs. 0.5–0.8 cm long), corolla tube sharply contracted (vs. gradually contracted).

Etymology: The name refers to the flowers, which are fasciculate on inflorescences of the new species.

Vernacular name: Cù Huā Qí Zhù Jù Tái (Chinese pronunciation); 簇花奇柱苣苔 (Chinese name).

Distribution and habitat: The new species only grows on the wet cliff in the valley and only occurs at the type locality, Jinping County, Yunnan province, China.

Yu-Min Shui, Jian-Yong Wu, Zhi-Yong Yu, Shi-Wei Guo, Li Chen, Fang Wen and Wen-Hong Chen. 2020. Deinostigma fasciculatum, A New Species of Gesneriaceae in Yunnan, China. In: Shui Y-M, Chen W-H, Ren M-X, Wen F, Hong X, Qiu Z-J, Wei Y-G, Kang M (Eds) Taxonomy of Gesneriaceae in China and Vietnam. PhytoKeys. 157: 199-206. DOI: 10.3897/phytokeys.157.32683

Didymocarpus lobulatus F. Wen, Xin Hong & W.Y. Xie

in Xie, ... et Wen, 2020.

DOI: 10.3897/phytokeys.157.30349

Abstract

Didymocarpus lobulatus, a new species endemic to Zhejiang province, eastern China, is described and illustrated with photographs. The new species is morphologically similar to D. heucherifolius, D. cortusifolius and D. salviiflorus in leaf morphology, but can be easily distinguished by a combination of characters, including the shape of bracts, calyx and calyx lobes.

Keywords: Flora of Zhejiang, new taxon, taxonomy, Didymocarpus sect. Heteroboea

Figure 1. Didymocarpus lobulatus
A Habit B Flower in top view C Opened corolla, showing stamens and staminodes D Fertile stamens and anthers E Pistil, disc and stigma.

Figure 2. Didymocarpus lobulatus F. Wen, Xin Hong & W.Y. Xie.
A Habitat B Vegetative part of plants C Habitat in flowering D Frontal view of corolla E Top view of cyme, showing actinomorphic calyx F Opened corolla for showing stamens, pistil and calyx lobes G Adaxial surface view of leaf blade H Abaxial surface view of leaf blade.

Didymocarpus lobulatus F. Wen, Xin Hong & W.Y. Xie, sp. nov.

Diagnosis: Didymocarpus lobulatus is similar to D. heucherifolius Handel-Mazzetti and D. salviiflorus W.Y. Chun in having a similar zygomorphic corolla and pink to pinkish-purple funnel-shaped to tubular corolla tube, but can be distinguished from the former by its densely eglandular and glandular pubescent peduncle (vs. villous in D. heucherifolius), bracts subulate to subulate-triangular (vs. elliptic) and margin sparsely crenate from the middle (vs. entire), calyx shallowly 5-lobed to or lobed about two-thirds of the calyx length from the base (vs. 5-lobed to the base). From D. salviiflorus, it differs by having subulate to subulate-triangular bracts (vs. semi-orbicular in D. salviiflorus), calyx lobes triangular but non-overlapping (vs. depressed oblong, overlapping at margin) and in size in ca. 5 × 2.5 mm (vs. 2–2.2 × 4–4.5 mm). It is also similar to D. cortusifolius in the shape and size of the leaves, but can be easily distinguished by having bracts subulate to subulate-triangular (vs. ovate to elliptic in D. cortusifolius), bract margin sparsely crenate from the middle (vs. entire), larger calyx lobes ca. 5 × 2.5 mm (vs. 1–3 × ca. 2 mm) and lobes’ margin nearly entire to entire (vs. denticulate), corolla white (vs. corolla pink to dark pink), fertile stamens adnate to corolla ca. 5 mm from base (vs. adnate to corolla 10–14 mm from base), filaments sparsely brownish glandular-puberulous (vs. glabrous).

Etymology: The specific epithet is derived from the shallowly 5-lobed calyx.

Distribution and habitat: Didymocarpus lobulatus is locally abundant and endemic to a narrow area in eastern China, surrounding the type locality. This species grows on moist shady cliffs of sandy shale hills, at an elevation of 223 m a.s.l. in type locality. The average temperature is 16.4 °C, the average annual precipitation has been calculated as ca. 1,446.8 mm. The forest is a subtropical monsoon climate evergreen broad-leaved forest. Flowering in May. Another population growing in the Danxia landscape of Chuanyanshijiufeng, Xinchang County, Shaoxing city, not far away from the type locality, was discovered by the first author in 2016. The two places are about 40 kilometres apart.

Figure 3. A Didymocarpus lobulatus B D. heucherifolius C D. cortusifolius D D. salviiflorus
1 Habitats 2 The frontal view of corolla and cyme 3 Cyme and bracts 4 calyx lobes.

Wen-Yuan Xie, Jia-Jun Zhou, Xin Hong and Fang Wen. 2020. Didymocarpus lobulatus (Gesneriaceae), A New Species from Zhejiang Province, East China. In: Shui Y-M, Chen W-H, Ren M-X, Wen F, Hong X, Qiu Z-J, Wei Y-G, Kang M (Eds) Taxonomy of Gesneriaceae in China and Vietnam. PhytoKeys. 157: 199-206. DOI: 10.3897/phytokeys.157.30349

Atheris hetfieldi

Ceríaco, Marques & Bauer, 2020

Hetfield’s Bush Viper || DOI: 10.11646/zootaxa.4838.4.9

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Illustration by Arthur C. Wandeur

Abstract
Two species of Bush Vipers, genus Atheris Cope, 1862, have been reported for the continental island of Bioko, Gulf of Guinea – Atheris squamigera and a putatively undescribed species, morphologically similar to Atheris chlorechis. The latter was only known from one specimen collected in the early 1900s and its taxonomic identity has never been fully resolved. Based on recently collected specimens of Atheris from Bioko Island, we reviewed all the available data and specimens through detailed morphological comparisons against all known species of the genus. Our results confirm that Atheris squamigera is present in Bioko Island, and that the second taxon is a new species, which we describe here as Atheris hetfieldi sp. nov. The newly described species presents a series of morphological characters that clearly differentiate it from all of its congeners. The description of this new species is the first of a new snake species from Bioko in more than 100 years and this species is the only snake currently recognized as endemic to the island.

Keywords: Reptilia, Taxonomy, snakes, morphology, endemic species, island biology

...

Morphological data lead us to recognize two species of Atheris in Bioko Island

– Atheris squamigera and a new species which we describe as new.

...

Atheris hetfieldi sp. nov.

Atheris squamigera squamigera [part]: Capocaccia (1961: 304)

Atheris chlorechis: Mertens (1964: 235)

Etymology. The species is named after James A. Hetfield (1963–present), lead vocalist and guitarist of the heavy metal band Metallica, for the inspiration, endurance and sanity that his music provides to the authors while roaming the academic world. The name is here applied as a genitive masculine. We propose the English common name of Hetfield’s Bush Viper.

Luis M. P. Ceríaco, Mariana P. Marques and Aaron. M. Bauer. 2020. The Bush Vipers, Genus Atheris Cope, 1862 (Squamata: Viperidae) of Bioko Island, Gulf of Guinea, with the Description of A New Species. Zootaxa. 4838(4); 581–593. DOI: 10.11646/zootaxa.4838.4.9

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Colobaea sp.

in Bratt, Knutson, et al. 2020.

DOI: 10.11646/zootaxa.4840.1.1

Abstract

All reared larvae of flies of the genus Colobaea Zetterstedt, 1837 (Diptera: Sciomyzidae), which comprises 15 valid species, kill and consume freshwater nonoperculate pulmonate snails. New data are presented on the geographic distribution, biology, morphology of immature stages, and classification of Colobaea. Life cycle information is provided from field data and laboratory rearings for four of the 11 Palearctic species—C. bifasciella (Fallén), C. deemingi Knutson & Bratt n.sp., C. pectoralis (Zetterstedt), and C. punctata (Lundbeck)—and for one of the three Nearctic species, C. americana Steyskal. Colobaea bifasciella is shown to be one of the most highly specialized parasitoid Sciomyzini, laying eggs on shells of Galba truncatula (O.F. Müller) and Stagnicola palustris (O.F. Müller) in temporary, intermittent, or vernal semiterrestrial situations. Each larva feeds in only one host snail, which is not killed until shortly before the larva completes development. Puparia are strongly modified to fit tightly within the shell of the host. The other reared species are shown to be less specialized than C. bifasciella, with eggs being laid upon vegetation, the larvae behaving as parasitoids-predators-saprophages of exposed aquatic snails, and the puparia of all four species being adapted to a lesser degree than C. bifasciella to fitting within the shell of the host snail. In nature, C. americana attacks Gyraulus parvus (Say) and Physa Draparnaud sp.; C. pectoralis attacks Anisus vortex (L.) and Bathyomphalus contortus (L.); and C. punctata attacks Gyraulus albus O.F. Müller, Lymnaea “peregra,” Planorbarius corneus (L.), and Planorbis planorbis (L.). In the laboratory, these species also attacked and consumed other freshwater nonoperculate snails; C. deemingi was reared on Gyraulus intermixtus (Mousson) and Radix gedrosiana (Say), and an adult fly of the Palearctic C. distincta (Meigen) emerged from a puparium found in the shell of Anisus spirorbis (L.) collected in nature. Described and figured are eggs, larvae of all three instars, and puparia of the five laboratory-reared species. To provide perspective on features of Colobaea, diagnostic features are summarized of the immature stages of the Sciomyzini and the suprageneric categories of Sciomyzidae. The biogeography of the tribe Sciomyzini is presented, along with details of known geographical distribution. The classification and phylogenetic position of Colobaea are discussed. Included are a checklist of all known taxa of Colobaea, maps of geographic distribution, and a key to adults of the 15 valid species.

Keywords: Diptera, Acalyptratae, Sciomyzoidea, snails, life cycles, morphology, immature stages, taxonomy, nomenclature, phylogenetic relationships

Albertus D. Bratt, Lloyd V. Knutson, William L. Murphy and Anthony A. Daniels. 2020. Biology, Immature Stages, and Systematics of Snail-killing Flies of the Genus Colobaea (Diptera: Sciomyzidae), with Overviews of Aspects of the Tribe Sciomyzini. Zootaxa. 4840(1); 1–64. DOI: 10.11646/zootaxa.4840.1.1

Anchylorhynchus sp.

in Medeiros & Vanin, 2020.

DOI: 10.11646/zootaxa.4839.1.1

Abstract

Anchylorhynchus Schoenherr is a genus of palm-associated weevils currently including 22 described species in the Neotropics. These weevils engage in brood pollination interactions with species in at least four genera of palms (Arecaceae), representing an emerging system for the study of mutualisms. Here we revise the taxonomy of Anchylorhynchus and propose the first phylogenetic hypothesis for the group, based on morphology. Anchylorhynchus chrysomeloides sp. nov., Anchylorhynchus goiano sp. nov., Anchylorhynchus imitator sp. nov., Anchylorhynchus latipes sp. nov., Anchylorhynchus multisquamis sp. nov. and Anchylorhynchus rectus sp. nov. are described, Anchylorhynchus gottsbergerorum Vanin is a new junior subjective synonym of Anchylorhynchus bicarinatus O’Brien, and Anchylorhynchus eriospathae Bondar and Anchylorhynchus pictipennis Hustache are new junior subjective synonyms of Anchylorhynchus tremolerasi Hustache, resulting in 25 valid species for the genus. We provide genus and species descriptions with a new dichotomous key to the species and updated information on geography and host associations based on museum records and extensive new collections. We also produce a new morphological matrix with 113 characters, 11 of them based on measurements while accounting for allometry, correlation and power to delimit groups. The software pipeline to produce these characters was encoded in a graphical user interface named DiscretzR, made available here. Analysis of this matrix under parsimony and Bayesian inference resulted in strong support for the monophyly of Anchylorhynchus, but weaker support for most clades within the genus. Reconstruction of the host plant associations indicates that the common ancestor of Anchylorhynchus visited flowers of Syagrus Mart., with later transitions to other genera restricted to one of the major clades of Anchylorhynchus. This taxonomic revision with the first phylogeny for the group provides a foundation for future evolutionary studies in the genus.

Keywords: Coleoptera, systematics, morphology, taxonomic revision, Coleoptera, Curculionidae, Arecaceae, Syagrus, Butia, Oenocarpus, Euterpe

Bruno A. S. de Medeiros and Sergio A. Vanin. 2020. Systematic Revision and Morphological Phylogenetic Analysis of Anchylorhynchus Schoenherr, 1836 (Coleoptera, Curculionidae: Derelomini). Zootaxa. 4839(1); 1-98. DOI: 10.11646/zootaxa.4839.1.1

Epipompilus namadgi

Yuan & Rodriguez, 2020

DOI: 10.11646/zootaxa.4743.4.7

twitter.com/juanita_rodr

Abstract
Three new species are added to the genus Epipompilus (Hymenoptera: Pompilidae) in Australia. Epipompilus mirabundus sp. nov., E. taree sp. nov., and E. namadgi sp. nov. are described and illustrated. A key to males of Epipompilus is provided. A novel association of Epipompilus and Sceliphron formosum (Hymenoptera: Sphecidae) is also documented. The larva of E. mirabundus sp. nov. was found sharing single nest cell with a Sceliphron larva; this association could be the result of a parasitised spider being brought back to the nest by the Sceliphron adult.

Keywords: Hymenoptera, Spider wasps, mud dauber wasps, Epipompilus, Sceliphron formosum

Epipompilus mirabundus Yuan & Rodriguez, sp. nov.

Distribution. Australia: Australian Capital Territory.

Etymology. The epithet originates from the Latin mirabundus, which means astonishment, and is based on the unusual way the specimen was found in its larval stage developing and sharing a cell with a Sceliphron formosum larva.

Remarks. The cocoon was found in the nest of Sceliphron formosum (Hymenoptera: Sphecidae) (Figs. 1D, 3). A male adult was reared and emerged in the lab.

Epipompilus taree Yuan & Rodriguez, sp. nov.

Distribution. Australia: New South Wales.

Etymology. The epithet, placed as a noun in apposition, comes from the city close to where the specimens were collected.

Epipompilus namadgi Yuan & Rodriguez, sp. nov.

Distribution. Australia: Australian Capital Territory.

Etymology. The epithet, placed as a noun in apposition, comes from the National Park where the type specimen was collected.

David Yuan and Juanita Rodriguez. 2020. Three New Species of Epipompilus Kohl (Hymenoptera, Pompilidae, Pepsinae) from Australia. Zootaxa. 4743(4); 575–584. DOI: 10.11646/zootaxa.4743.4.7

twitter.com/juanita_rodr/status/1232942310620446721

bushblitz.org.au/a-new-wasp-species-with-alzheimers-fighting-venom-may-be-lost-to-science-due-to-bushfires

@puppyyuan #spiderwasp @BushBlitz2 #australianbushfires #namadgi

Epipogium taiwanense T.C.Hsu

in Wu, Liu, Tang, ... et Peng, 2020.

DOI: 10.11646/phytotaxa.453.1.4

Abstract

Epipogium taiwanense, is published in the book of Illustrated Flora of Taiwan in 2018. However, here only the specimen and simply descriptions were existed on which the protologue was based. Plant and flowers characteristics were not described in enough detail in the original description. Fortunately, it was rediscovered from Yunnan and Sichuan Province, China. It is morphological similar to E. aphyllum, but it can be easily distinguished from the latter by having large size flower bract (2.0 × 1.0 cm, significantly longer than pedicel and ovary), epichile of labellum concave with 1 longitudinal ridge on mid-lobe and the spur strongly recurve in the apex connecting with bottom of mid-lobe. A preliminary risk-of-extinction assessment, according to the IUCN Red List Categories and Criteria, is given for the newly recorded species.

Keywords: Monocots, Epipogium, Morphological taxonomy, ghost orchid, Epidendrea, Orchidaceae

Xun-Feng Wu, Qiang Liu, Ying Tang, Kuan-Bo Chi, Yu-Qian Wang, Xin Pang, Yao-Hua Huang and Pei-Hao Peng. 2020. Epipogium taiwanense, A detail Morphological Description and Additional New Record to Mainland China. Phytotaxa. 453(1); 43–48. DOI: 10.11646/phytotaxa.453.1.4

臺灣上鬚蘭

Jurassic belemnite jaws: Acrocoelites conoideus & Hibolithes semisulcatus.

in Klug, Etter, et al., 2020.

DOI: 10.1186/s13358-020-00207-7

Reconstructions by Kenneth De Baets.

twitter.com/djbirddanerd

Abstract
Although belemnite rostra can be quite abundant in Jurassic and Cretaceous strata, the record of belemnite jaws was limited to a few specimens from Germany and Russia. Here, we describe and figure three cephalopod jaws from the Middle Jurassic Opalinus Clay of northern Switzerland. Although flattened, the carbonaceous fossils display enough morphological information to rule out an ammonoid, nautiloid or octobrachian origin of the two larger jaws. Their similarities to belemnite jaws from Germany and Russia conforms with our interpretation of these specimens as belemnite jaws. Based on their rather large size, we tentatively assign these two jaws to the megateuthidid Acrocoelites conoideus. The third jaw is a rather small upper jaw of an ammonoid. Since Leioceras opalinum is by far the most common ammonite in this unit in northern Switzerland, we tentatively suggest that the upper jaw belongs to this species.

Keywords: Cephalopoda, Megateuthididae, Graphoceratidae, Mouthparts, Body size, Opalinus clay

Reconstructions of Jurassic belemnite jaws.
a–c Acrocoelites conoideusfrom the Swiss Aalenian.
d–f Hibolithes semisulcatusfrom the German Kimmeridgian, modified after Klug et al. (2010b); the inner lamella of the lower jaw was enlarged

Christian Klug, Walter Etter, René Hoffmann, Dirk Fuchs and Kenneth De Baets. 2020. Jaws of A Large Belemnite and An Ammonite from the Aalenian (Middle Jurassic) of Switzerland. Swiss Journal of Palaeontology. 139, 4. DOI: 10.1186/s13358-020-00207-7

twitter.com/djbirddanerd/status/1300415883744022530

Neolithodes indicus

Padate, Cubelio & Takeda, 2020

DOI: 10.11646/zootaxa.4845.1.5

Abstract

A new species of the deep-water king-crab genus Neolithodes A. Milne-Edwards & Bouvier, 1894 (Anomura: Lithodidae) is described herein from the southeastern Arabian Sea (1064–1338 m depth). Neolithodes indicus sp. nov. shows morphological proximity with three congeneric species, N. brodiei Dawson & Yaldwyn, 1970, N. flindersi Ahyong, 2010a and N. nipponensis Sakai, 1971 in bearing less numerous secondary spines on the carapace, the dactylus of the chelipeds with a convex dorsal margin and dorsoventrally compressed meri of the pereopods 2–4. The new species can be easily distinguished from these congeners in possessing the less densely spinulose carapace and appendages, short rostrum, simple antennal scaphocerite, the absence of large spines on the flexor margins of the pereopods 2–3 meri, and a stouter pereopod 4 propodus.

Keywords: Crustacea, Lithodidae, Neolithodes, taxonomy, India

Vinay P. Padate, Sherine Sonia Cubelio and Masatsune Takeda. 2020. Description of A New Species of Deep-water King-crab (Crustacea: Decapoda: Anomura) from the southeastern Arabian Sea. Zootaxa. 4845(1); 71–82. DOI: 10.11646/zootaxa.4845.1.5

Sciodaphyllum zarucchii M. M. Mora, Lowry, Idárraga, Jiménez-Mont. & G. M. Plunkett

in Mora, Lowry, Plunkett, ... et Raven, 2020.

DOI: 10.3417/2020540

Abstract

Sciodaphyllum zarucchii M. M. Mora, Lowry, Idárraga, Jiménez-Mont. & G. M. Plunkett (Araliaceae) is described as a new species in honor of James L. Zarucchi (1952–2019). It occurs in humid premontane and montane forests on the western slope of the Cordillera Occidental in the department of Antioquia, Colombia, where it is known from only two localities, one of which is highly threatened by forest clearing. A risk of extinction assessment using the IUCN Red List criteria reveals that S. zarucchii is Endangered.

KEYWORDS: Andes, Araliaceae, conservation status, Neotropics, new species, Schefflera, Sciodaphyllum

Figure 2. Holotype of Sciodaphyllum zarucchii M. M. Mora, Lowry, Idárraga, Jiménez-Mont. & G. M. Plunkett (Zarucchi, Brandt & Castaño 5634, MO).

Figure 1. Photos of Sciodaphyllum zarucchii M. M. Mora, Lowry, Idárraga, Jiménez-Mont. & G. M. Plunkett.
-A. Branch with young infructescence. -B. Leaf (note absence of stipular ligule at the base of each of the three petioles, indicated by arrows). -C. Inflorescence branch with flower buds (note yellowing stipular ligule about to abscise from petiole base of the uppermost leaf, indicated by an arrow). -D. Heads with immature fruits.

From Jiménez et al. 2295 (HUA). Photographs by J. Jiménez-Montoya.

Sciodaphyllum zarucchii M. M. Mora, Lowry, Idárraga, Jiménez-Mont. & G. M. Plunkett

Etymology. This tree is named in honor of James L. Zarucchi (1952–2019), who joined the staff of the Missouri Botanical Garden in 1983. Both in the course of his doctoral studies at Harvard University (Ph.D.,1982) and for his first 13 years at the Garden, Jim concentrated on Latin American plants. He notably collected and encouraged studies of the flora of the Andes, especially in the departments of Antioquia and Vaupes, Colombia, coincidentally describing the genus Quiotania Zarucchi (Apocynaceae), its name an anagram for that of the Colombian Department of Antioquia. In 1996, Jim was named Managing Editor of the Flora of North America project, making an extensive contribution of great importance in that role for the remaining 23 years of his life (Ulloa et al., 2019).

M. Marcela Mora, Porter P. Lowry II, Gregory M. Plunkett, Álvaro Idárraga-Piedrahíta, Jáider Jiménez-Montoya and Peter H. Raven. 2020. Studies in Neotropical Araliaceae. V. Sciodaphyllum zarucchii (Araliaceae), A New Species from Antioquia, Colombia, Honoring James L. Zarucchi (1952–2019). Novon: A Journal for Botanical Nomenclature. 28(2); 94-99. DOI: 10.3417/2020540

Researchgate.net/publication/341335409_Sciodaphyllum_zarucchii_a_New_Species_from_Antioquia_Colombia

in Sudasinghe, Pethiyagoda, Raghavan, et al., 2020.

DOI: 10.1111/zsc.12445

facebook.com/HiranyaSudasinghe

Abstract

The South and South‐East Asian freshwater fish genus Systomus (Cyprinidae) comprises 17 valid species. Six nominal species, including three endemics, have been reported from Sri Lanka, a continental island separated from India by a shallow‐shelf sea. The species diversity of Systomus on the island has until now not been assessed; neither has an evaluation been made of their phylogenetic history. Here, based on an analysis of the nuclear recombination activating protein 1 (rag1), and mitochondrial cytochrome c oxidase subunit 1 (cox1) and cytochrome b (cytb) gene markers, and a morphological examination of 143 specimens from 49 locations in Sri Lanka, we reassess the diversity of Systomus on the island and analyse patterns of their evolution and biogeography. Divergence‐time estimates, based on a substitution rate calibration, date the basal split between Systomus and its sister group, the Afrotropical small barbs, to 30.0 Ma (95% highest posterior density: 25.4–35.2 Ma). The species of Systomus belong to two distinct clades. The first includes the Sri Lankan endemics S. asoka, S. martenstyni and S. pleurotaenia, which comprise an insular diversification following the immigration of a common ancestor during the Oligocene. The second, which includes the remaining species of Indian, Sri Lankan and South‐East Asian Systomus, has a crown age dating to the Late Miocene. Morphological and molecular species delimitation analyses failed to validate the two nominal species, S. spilurus and S. timbiri, previously reported from Sri Lanka: both are considered synonyms of S. sarana, as are the nomina S. chryseus, S. chrysopoma, S. laticeps, S. rufus, S. pinnauratus and S. subnasutus. Four genetically and geographically discrete lineages of S. sarana occur in the island, and three in India. Molecular species delimitation analysis suggests these all belong to a single species, S. sarana. The genetically distinct Sri Lankan populations of S. sarana result from Plio‐Pleistocene dispersal or vicariance events between India and Sri Lanka—as a result of emergence and inundation of the now submerged isthmus connecting the two landmasses—as well as autochthonous insular diversification.

Keywords: diversification, molecular dating, molecular systematics, phylogeography, Smiliogastrinae, taxonomy

Hiranya Sudasinghe, Rohan Pethiyagoda, Rajeev Raghavan, Neelesh Dahanukar, Lukas Rüber and Madhava Meegaskumbura. 2020. Diversity, Phylogeny and Biogeography of Systomus (Teleostei, Cyprinidae) in Sri Lanka. Zoologica Scripta. DOI: 10.1111/zsc.12445

facebook.com/HiranyaSudasinghe/posts/10221031740276095

Paracobitis salihae

Kaya, Turann, Kalayci, Bayçelebi & Freyhof, 2020

DOI: 10.11646/zootaxa.4838.4.6

twitter.com/Jorg_Freyhof

Abstract

Paracobitis salihae, new species, from the Göksu River in the western upper Euphrates drainage, is distinguished from other Paracobitis species by possessing a truncate caudal-fin, and a dark-brown vermiculate or marbled colour pattern. It is also characterised by 19 variable nucleotide substitutions, three diagnostic nucleotides and 3.6% minimum K2P distance compared to geographically adjacent and related P. zabgawraensis. Paracobitis salihae might be Critically Endangered and is threatened by dam constructions.

Keywords: Pisces, Freshwater fish, Taxonomy, Cytochrome oxidase I, Turkey

Paracobitis salihae, FFR 3657, holotype, 66 mm SL; Turkey: Göksu River.

Paracobitis salihae, new species

Distribution. Only known from the Göksu River, a tributary of the upper Euphrates (Figs. 5–6).

Etymology. Paracobitis salihae is named in honour of the Saliha Kaya (1939-2015), mother of the first author. A noun in the genitive, indeclinable.

Cüneyt Kaya, Davut Turann, Gökhan Kalayci, Esra Bayçelebi and Jörg Freyhof. 2020. The Westernmost known Population of Paracobitis (Teleostei, Nemacheilidae), with the Description of A New Species from the Euphrates River in southern Anatolia. Zootaxa. 4838(4); 525–534. DOI: 10.11646/zootaxa.4838.4.6

twitter.com/Jorg_Freyhof/status/1299287819534430215

Dixonius pawangkhananti

Pauwels, Chomngam, Larsen & Sumontha, 2020

จิ้งจกดินชะอำ || DOI: 10.11646/zootaxa.4845.1.7

facebook.com/MontriSumontha

facebook.com/ItalianGekkoAssociation

Abstract

We describe Dixonius pawangkhananti sp. nov. from coastal limestone hills in Cha-am District, Phetchaburi Province, peninsular Thailand. The new species differs from all currently recognized Dixonius by the following combination of morphological characters and pattern: maximal known snout-vent length of 42.6 mm; 16 longitudinal rows of dorsal tubercles; 30 to 32 paravertebral scales; 16 longitudinal rows of ventral scales across the abdomen; six precloacal pores in males, no pores in females; a marked canthal stripe; and a sexually dimorphic dorsal pattern consisting of bands (males) or blotches (females). This description brings the number of Dixonius species to ten, with four species endemic to Thailand.

Keywords: Gekkonidae, Thai-Malay Peninsula, Dixonius pawangkhananti sp. nov.

Live adult female (above) and male Dixonius pawangkhananti sp. nov. in ... Phetchaburi Province, Thailand. Individuals not collected. Photo. by M. Sumontha.

Dixonius pawangkhananti sp. nov.

Etymology. The specific epithet honors the Thai zoologist Parinya Pawangkhanant (Agriculture University of Phayao, Phayao, and Rabbit in the Moon Foundation, Ratchaburi), for his contributions to the herpetology of Thailand, and who is one of the collectors of the type-series. We suggest the following common names: Djing-djok din Cha-am (Thai: จิ้งจกดินชะอำ), Cha-am leaf-toed gecko (English), Dixonius de Cha-am (French), Cha-am Blattfingergecko (German).

Biotope where Dixonius pawangkhananti sp. nov. lives in ... Forest Park, Phetchaburi Province, Thailand. Photo. by P. Pawangkhanant.

Olivier S. G. Pauwels, Nirut Chomngam, Henning Larsen and Montri Sumontha. 2020. A New Limestone-dwelling Leaf-toed Gecko (Gekkonidae: Dixonius) from Coastal Hills in Cha-am, peninsular Thailand. Zootaxa. 4845(1); 97–108. DOI: 10.11646/zootaxa.4845.1.7

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Nabangana koreana Byun, 2020

DOI: 10.11646/zootaxa.4845.2.9

Abstract

Nabangana gen. nov. is described and illustrated from Korea. The new genus and its type species Nabangana koreana sp. nov. represents the first record of the tortricid tribe Chlidanotini from Korea.

Key words: Chlidanotinae, Korea, Nabangana, new taxa

Morphological features of adults of Nabangana koreana sp. nov.
1, adult, paratype; 2, frontal view of head; 3, lateral view of head; 4, wing venation.

Genitalia of Nabangana koreana sp. nov.
5, male genitalia, holotype (genitalia slide number 854); 6, female genitalia, paratype (genitalia slide number 3464). Scale bar: 0.5mm.

Tribe Chlidanotini Meyrick, 1906

Type genus: Chlidanota Meyrick, 1906

Journal of the Bombay Natural History Society, 17 (2): 412.

Genus Nabangana gen. nov.

Type species: Nabangana koreana sp. nov.

Diagnosis. The new genus is somewhat similar in appearance to members of the tribe Enarmoniini, but it can be easily distinguished by the concaved termen beneath the apex in the forewing, the presence of hami and a subtriangular lobe on the valva in male genitalia, and a signum bearing numerous long spines in female genitalia. Also, the new genus is similar to the related genera, Trymalitis Meyrick, 1905, Caenognosis Walsingharn, 1900, Leurogyia Common, 1965, and Daulocnema Common, 1965, but it is different from them by the wing venation as written in the Discussion. The male genitalia (Figs 5, 5a) are similar to those of Leurogyia peristictum Common, 1965, but can be distinguished by the rather narrow uncus, with sub-acute apex; the stick-shaped hami with broadened apex; the rather rounded valva, curved at middle of ventral margin with a subtriangular lobe medially. Aedeagus nearly straight with bifid apex, cornuti absent. The female genitalia (Fig. 6) are in general similar to those of allied genera, but are characteristic especially by the sclerotized ductus bursae near the entrance, and the signum with a bundle of spines, and the accessory sac originating from the head of the signum.

Distribution. The genus occurs only in Korea.

Etymology. The species name of the new genus is derived from the common name for “moth” in Korea (Nabang).

Nabangana koreana sp. nov.

Diagnosis. The new species is similar to members of the allied genera, Caenognosis incisa Walsingham, 1900, Leurogyia peristictum Common, 1965, and Daulocnema epicharis Common, 1965 in wing pattern, color, and the shape of the forewing termen, but it can be distinguished by the elongate uncus, the golf club-shaped hami, the curved costa of the valva, and a nearly straight aedeagus. In wing venation, the new species is quite similar to Leurogyia peristictum, with the forked R4 and R5 near the apex in the forewing; however, in the latter, the veins are positioned near the apex at the costa and beneath the apex at the termen, respectively, whereas in N. koreana R4 and R5 are positioned together near the apex at the costa.

Etymology. The species name is derived from the type locality (Korea).

Bong-Kyu Byun. 2020. First Report of the Tribe Chlidanotini (Lepidoptera: Tortricidae) from Korea, with the Description of A New Genus and Species. Zootaxa. 4845(2); 283–287. DOI: 10.11646/zootaxa.4845.2.9

[Herpetology • 2020] Pristimantis zorro • Out of the Blue: A New Rain Frog Species of the Genus Pristimantis (Anura: Craugastoridae) from the northern Cordillera Central in Colombia

[Herpetology • 2020] Cyrtodactylus amphiptraeus • A New Species Cyrtodactylus Gray (Squamata: Gekkonidae) from western Thailand and the Phylogenetic Placement of C. inthanon and C. doisuthep

[Botany • 2020] Impatiens dindigulensis (Balsaminaceae) • A New Species from Kodaikkanal Wildlife Sanctuary, India

[Ichthyology • 2020] Betta nuluhon • A New Species of Fighting Fish (Teleostei: Osphronemidae) from western Sabah, Malaysia

[Herpetology • 2020] Cnemaspis rishivalleyensis & Hemidactylus rishivalleyensis • Geckos in the Granite: Two New Geckos (Squamata: Gekkonidae) from Rocky, Scrub Habitats in Rishi Valley, Andhra Pradesh, India

[Crustacea • 2020] Turleania rubriguttatus • A New Species of Pagurid Hermit Crab (Decapoda: Anomura: Paguroidea) from Shallow Water in Japan, with Notes on T. senticosa (McLaughlin & Haig, 1996)

[Botany • 2020] Deinostigma fasciculatum (Gesneriaceae) • A New Species from Yunnan, China

[Botany • 2020] Didymocarpus lobulatus (Gesneriaceae) • A New Species from Zhejiang Province, East China

[Herpetology • 2020] Atheris hetfieldi • The Bush Vipers, Genus Atheris Cope, 1862 (Squamata: Viperidae) of Bioko Island, Gulf of Guinea, with the Description of A New Species

[Entomology • 2020] Biology, Immature Stages, and Systematics of Snail-killing Flies of the Genus Colobaea (Diptera: Sciomyzidae), with Overviews of Aspects of the Tribe Sciomyzini

[Entomology • 2020] Systematic Revision and Morphological Phylogenetic Analysis of Anchylorhynchus Schoenherr, 1836 (Coleoptera, Curculionidae: Derelomini)

[Entomology • 2020] Three New Species of Epipompilus Kohl (Hymenoptera, Pompilidae, Pepsinae) from Australia

[Botany • 2020] Epipogium taiwanense (Orchidaceae) • A detail Morphological Description and Additional New Record to Mainland China

[Paleontology • 2020] Jaws of A Large Belemnite and An Ammonite from the Aalenian (Middle Jurassic) of Switzerland

[Crustacea • 2020] Neolithodes indicus • A New Species of Deep-water King-crab (Decapoda: Anomura) from the southeastern Arabian Sea

[Botany • 2020] Sciodaphyllum zarucchii (Araliaceae) • A New Species from Antioquia, Colombia [Studies in Neotropical Araliaceae. V]

[Ichthyology • 2020] Diversity, Phylogeny and Biogeography of Systomus (Teleostei, Cyprinidae) in Sri Lanka

[Ichthyology • 2020] Paracobitis salihae • The Westernmost known Population of Paracobitis (Teleostei, Nemacheilidae), with the Description of A New Species from the Euphrates River in southern Anatolia

[Herpetology • 2020] Dixonius pawangkhananti จิ้งจกดินชะอำ • A New Limestone-dwelling Leaf-toed Gecko (Gekkonidae: Dixonius) from Coastal Hills in Cha-am, peninsular Thailand

[Entomology • 2020] Nabangana koreana • First Report of the Tribe Chlidanotini (Lepidoptera: Tortricidae) from Korea, with the Description of A New Genus and Species